COCHLEATA controls leaf size and secondary inflorescence architecture via negative regulation of UNIFOLIATA (LEAFY ortholog) gene in garden pea Pisum sativum
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Title |
COCHLEATA controls leaf size and secondary inflorescence architecture via negative regulation of UNIFOLIATA (LEAFY ortholog) gene in garden pea Pisum sativum
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Creator |
Sharma, Vishakha
Chaudhary, Swati Kumar, Arvind Kumar, Sushil |
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Subject |
Bracteole differentiation
COCHLEATA gene garden pea leaf differentiation LEAFY ortholog leaf size Pisum sativum secondary inflorescence stub histology UNIFOLIATA gene UNIFOLIATA repression UNIFOLIATA-TENDRILLED ACACIA gene |
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Description |
UNIFOLIATA [(UNI) or UNIFOLIATA-TENDRILLED ACACIA (UNI-TAC)] expression is known to be negatively regulated by COCHLEATA (COCH) in the differentiating stipules and flowers of Pisum sativum. In this study, additional roles of UNI and COCH in P. sativum were investigated. Comparative phenotyping revealed pleiotropic differences between COCH (UNI-TAC and uni-tac) and coch (UNI-TAC and uni-tac) genotypes of common genetic background. Secondary inflorescences were bracteole-less and bracteolated in COCH and coch genotypes, respectively. In comparison to the leaves and corresponding sub-organs and tissues produced on COCH plants, coch plants produced leaves of 1.5-fold higher biomass, 1.5-fold broader petioles and leaflets that were 1.8-fold larger in span and 1.2-fold dorso-ventrally thicker. coch leaflets possessed epidermal cells 1.3-fold larger in number and size, 1.4-fold larger spongy parenchyma cells and primary vascular bundles with 1.2-fold larger diameter. The transcript levels of UNI were at least 2-fold higher in coch leaves and secondary inflorescences than the corresponding COCH organs. It was concluded that COCH negatively regulated UNI in the differentiating leaves and secondary inflorescences and thereby controlled their sizes and/or structures. It was also surmised that COCH and UNI (LFY homolog) occur together widely in stipulate flowering plants.
We wish to acknowledge support of the Director, NIPGR, in terms of provision of research facilities and help provided by Vinod Kumar in field crop management. The research was funded by Indian National Science Academy and Council of Scientific and Industrial Research (CSIR, India) grants to SK and postgraduate fellowship grants from CSIR to SC and SK and from SKA Institution for Research, Education and Development to VS. We also wish to thank Susan Singer for the highly useful discussion via email and Anand Sarkar and Bithika Sharma for reading the manuscript. |
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Date |
2015-10-29T07:19:45Z
2015-10-29T07:19:45Z 2012 |
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Type |
Article
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Identifier |
Journal of Biosciences, 37(6): 1041-1059
0973-7138 http://www.ias.ac.in/jbiosci/dec2012/1041 http://172.16.0.77:8080/jspui/handle/123456789/280 |
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Language |
en_US
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Publisher |
Indian Academy of Sciences
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