Heterotic patterns of primary and secondary metabolites in the oilseed crop Brassica juncea
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Title |
Heterotic patterns of primary and secondary metabolites in the oilseed crop Brassica juncea
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Creator |
Bajpai, Prabodh K.
Reichelt, Michael Augustine, Rehna Gershenzon, Jonathan Bisht, Naveen C. |
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Subject |
Brassica juncea
oilseed crop primary and secondary metabolites Heterotic patterns |
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Description |
Accepted date: 11 March 2019
Heterosis refers to the superior performance of F1 hybrids over their respective parental inbred lines. Although the genetic and expression basis of heterosis have been previously investigated, the metabolic basis for this phenomenon is poorly understood. In a preliminary morphological study in Brassica juncea, we observed significant heterosis at the 50% flowering stage, wherein both the growth and reproduction of F1 reciprocal hybrids were greater than that of their parents. To identify the possible metabolic causes or consequences of this heterosis, we carried out targeted LC-MS analysis of 48 primary (amino acids and sugars) and secondary metabolites (phytohormones, glucosinolates, flavonoids, and phenolic esters) in five developmental tissues at 50% flowering in hybrids and inbred parents. Principal component analysis (PCA) of metabolites clearly separated inbred lines from their hybrids, particularly in the bud tissues. In general, secondary metabolites displayed more negative heterosis values in comparison to primary metabolites. The tested primary and secondary metabolites displayed both additive and non-additive modes of inheritance in F1 hybrids, wherein the number of metabolites showing an additive mode of inheritance were higher in buds and siliques (52.77–97.14%) compared to leaf tissues (47.37–80%). Partial least regression (PLS) analysis further showed that primary metabolites, in general, displayed higher association with morphological parameters in F1 hybrids. Overall, our results are consistent with a resource-cost model for heterosis in B. juncea, where metabolite allocation in hybrids appears to favor growth, at the expense of secondary metabolism. The work was supported by NIPGR core grant to NCB. PKB acknowledge the National Post-doctoral fellowship (NPDF grant no. PDF/2015/000391/LS) from SERB, DST (India). RA was funded with short-term research fellowship from NIPGR, India. NCB acknowledges the Max Planck-India mobility grant jointly funded by Max Planck Society (Germany) and DST (India). MR and JG acknowledge the funding by the Max Planck Society (Germany). We sincerely thank Dr. Payal Sanadhya (PS) for her help in drawing Fig. 4 and Dr. Arun Jagannath for language editing. Insightful suggestions from two anonymous reviewers are highly acknowledged. |
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Date |
2019-03-28T06:48:16Z
2019-03-28T06:48:16Z 2019 |
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Type |
Article
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Identifier |
Heredity, 123(3): 318-336
1365-2540 http://223.31.159.10:8080/jspui/handle/123456789/935 https://www.nature.com/articles/s41437-019-0213-3 https://doi.org/10.1038/s41437-019-0213-3 |
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Language |
en_US
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Format |
application/pdf
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Publisher |
Springer Nature
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